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The steps we need to take to solve for equilibrium in population genetics models follow the same general pattern regardless of what model we are analyzing. We emphasize that these models were developed to help us understand the genetic changes taking place in a population. The models shed light on nonintuitive processes and help quantify intuitive processes. The steps in the models can be outlined as follows: 1. Set up an algebraic model. 2. Calculate allelic frequency in the next generation, qn 1. 3. Calculate change in allelic frequency between generations, q. ^ 4. Calculate the equilibrium condition, q (q-hat), at q 0. 5. Determine, when feasible, if the equilibrium is stable.

For C# only, add the following using statement at the start of the code window: using Microsoft.VisualBasic;

Resize(id, 0, 0) Locate(id, ABS, 0, 0)

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in which pn is the increment of a alleles added by forward mutation, and qn is the loss of a alleles due to back mutation. Equation 20.1 takes into account not only the rate of forward mutation, , but also pn, the frequency of A alleles available to mutate. Similarly, the loss of a to A alleles is the product of both the rate of back mutation, , and the frequency of the a allele, qn. Equation 20.1 completes the second modeling step, derivation of an expression for qn 1, allelic frequency after one generation of mutation pressure. The third step is to derive an expression for the change in allelic frequency between two generations.This change ( q) is simply the difference between the allelic frequency at generation n 1 and the allelic frequency at generation n. Thus, for the a allele q qn

(20.2)

Next, if the argument loop contains the string value stop , then any currently playing sound is stopped by setting the innerHTML property of id to the empty string, thus removing any previously embedded sound player:

(20.3)

Note For C# only, you will also need to follow the steps in the Setting a Reference to the VB Namespace

if (loop == 'stop') O(id)innerHTML = ''

The next step in the model is to calculate the equilib^ rium condition q, or the allelic frequency when there is no change in allelic frequency from one generation to the next that is, when q (equation 20.3) is equal to zero: q Thus, pn Then, substituting (1 qn 1

(20.5)

Otherwise, the innerHTML property of id is assigned the correct HTML to embed a sound player and auto start the sound playing, looping it if loop contains the string value true , like this:

(20.4)

2. Take some time to explore the existing application controls and code. You will find some errors in the Task List because this code is not complete. Locate the comment in the code window that says "ADD CODE HERE!". 3. Directly under the line that says "Add Code Here", add code to sleep for 1 second (1,000 milliseconds). This is necessary because the line just above the comment has launched the Calculator accessory, and we need to make sure that it has had a chance to launch before sending keystrokes to it. System.Threading.Thread.Sleep(1000) You can type this line of code or you can write the code at the top of the file to import the System. Threading.Thread namespace, and then simply call the Sleep() method by itself: Sleep(1000).

or, by rearranging:

else O(id)innerHTML = InsVars("<embed src='#1' hidden='true' " + "autostart='true' loop='#2' />", file, loop)

(20.6)

Playing a sound is as easy as passing an empty container such as a div or span to the plug-in, along with the URL of the sound to play and, if required, the value true in the argument loop Here s an example that attaches the plug-in to the onmouseover events of four images:

And, since p q 1,

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